About the Authors

Ritobrato Datta

Affiliation Department of Neurology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jongho Lee

Affiliation Department of Radiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jeffrey Duda

Affiliation Department of Radiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Brian B. Avants

Affiliation Department of Radiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Charles H. Vite

Affiliation Section of Neurology, Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Ben Tseng

Affiliation Department of Neurology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

James C. Gee

Affiliation Department of Radiology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gustavo D. Aguirre

Affiliation Section of Ophthalmology, Department of Clinical Studies, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Geoffrey K. Aguirre

aguirreg@mail.med.upenn.edu

Affiliation Department of Neurology, School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JL GDA GKA. Performed the experiments: JL GDA GKA. Analyzed the data: RD JD BBA CHV BT JCG GKA. Contributed reagents/materials/analysis tools: BBA JCG. Wrote the paper: RD GKA.