About the Authors

Shigeyuki Yamamoto

Affiliations Department of Physiology, Osaka City University Graduate School of Medicine, Abeno-ku, Osaka, Japan, Central Research Laboratory, Hamamatsu Photonics KK, Hamakita, Shizuoka, Japan

Yasuomi Ouchi

Affiliation Molecular Imaging Frontier Research Center, Hamamatsu University School of Medicine, Hamamatsu, Japan

Daisaku Nakatsuka

Affiliation Department of Physiology, Osaka City University Graduate School of Medicine, Abeno-ku, Osaka, Japan

Tsuyoshi Tahara

Affiliation RIKEN Center for Molecular Imaging Science (CMIS), Kobe, Hyogo, Japan

Kei Mizuno

Affiliation RIKEN Center for Molecular Imaging Science (CMIS), Kobe, Hyogo, Japan

Seiki Tajima

Affiliation Department of Physiology, Osaka City University Graduate School of Medicine, Abeno-ku, Osaka, Japan

Hirotaka Onoe

Affiliation RIKEN Center for Molecular Imaging Science (CMIS), Kobe, Hyogo, Japan

Etsuji Yoshikawa

Affiliation Central Research Laboratory, Hamamatsu Photonics KK, Hamakita, Shizuoka, Japan

Hideo Tsukada

Affiliation Central Research Laboratory, Hamamatsu Photonics KK, Hamakita, Shizuoka, Japan

Masao Iwase

Affiliation Psychiatry, Department of Clinical Neuroscience, Osaka University Graduate School of Medicine, Suita, Japan

Kouzi Yamaguti

Affiliation Department of Physiology, Osaka City University Graduate School of Medicine, Abeno-ku, Osaka, Japan

Hirohiko Kuratsune

Affiliation Department of Health Sciences, Faculty of Health Sciences for Welfare, Kansai University of Welfare Sciences, Kashiwara, Japan

Yasuyoshi Watanabe

yywata@riken.jp

Affiliations Department of Physiology, Osaka City University Graduate School of Medicine, Abeno-ku, Osaka, Japan, RIKEN Center for Molecular Imaging Science (CMIS), Kobe, Hyogo, Japan

Competing Interests

The authors have read the journal’s policy and have the following conflicts: SY, EY and HT are employees of Hamamatsu Photonics KK. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: YW. Performed the experiments: SY YO DN TT ST EY HT MI KY HK. Analyzed the data: SY TT KM ST EY. Wrote the paper: SY YO KM HO YW.