About the Authors

Wentao Song

Contributed equally to this work with: Wentao Song, Renyi Pang

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, Weihai Vocational College, Department of Biological and Chemical Engineering, Weihai, China

Renyi Pang

Contributed equally to this work with: Wentao Song, Renyi Pang

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Yuze Niu

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, College of Fisheries and Life Science, Shanghai Ocean University, Shanghai, China

Fengtao Gao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Yongwei Zhao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Jing Zhang

Affiliations Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China, College of Fisheries and Life Science, Shanghai Ocean University, Shanghai, China

Jian Sun

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Changwei Shao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Xiaolin Liao

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Lei Wang

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Yongsheng Tian

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Songlin Chen

chensl@ysfri.ac.cn

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SC WS CS XL. Performed the experiments: WS RP YN FG YZ JZ JS. Analyzed the data: WS. Contributed reagents/materials/analysis tools: LW YT SC. Wrote the paper: WS SC.