About the Authors

Zhenhai Chen

Affiliation Department of Infectious Diseases, University of Georgia College of Veterinary Medicine, Athens, Georgia, United States of America

Pei Xu

Affiliations Department of Infectious Diseases, University of Georgia College of Veterinary Medicine, Athens, Georgia, United States of America, Intercollege Graduate Program in Cell and Developmental Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Gregory W. Salyards

Current address: California National Primate Research Center, University of California–Davis, Davis, California, United States of America

Affiliation University Research Animal Resources and the Department of Population Health, College of Veterinary Medicine, University of Georgia, Athens, Georgia, United States of America

Stephen B. Harvey

Affiliation University Research Animal Resources and the Department of Population Health, College of Veterinary Medicine, University of Georgia, Athens, Georgia, United States of America

Balazs Rada

Affiliation Department of Infectious Diseases, University of Georgia College of Veterinary Medicine, Athens, Georgia, United States of America

Zhen F. Fu

Affiliation Department of Pathology, University of Georgia College of Veterinary Medicine, Athens, Georgia, United States of America

Biao He

bhe@uga.edu

Affiliation Department of Infectious Diseases, University of Georgia College of Veterinary Medicine, Athens, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZC PX GWS SBH ZFF BH. Analyzed the data: ZC PX GWS SBH BR ZFF BH. Wrote the paper: ZC PX GWS SBH BH. Designed the experiments: ZC PX GWS SBH BR ZFF BH. Performed experiments: ZC PX GWS SBR BR.