About the Authors

Tim R. H. Read

tread@mshc.org.au

Affiliations Melbourne Sexual Health Centre, Alfred Health, Melbourne, Australia, School of Population Health, University of Melbourne, Melbourne, Australia

Jane S. Hocking

Affiliation Centre for Women’s Health, Gender and Society, University of Melbourne, Melbourne, Australia

Lenka A. Vodstrcil

Affiliations Melbourne Sexual Health Centre, Alfred Health, Melbourne, Australia, School of Population Health, University of Melbourne, Melbourne, Australia

Sepehr N. Tabrizi

Affiliation Dept of Microbiology and Infectious Diseases, Royal Women’s Hospital, Department of Obstetrics and Gynaecology, University of Melbourne, Murdoch Children’s Research Institute, Melbourne, Australia

Michael J. McCullough

Affiliation Melbourne Dental School, University of Melbourne, Melbourne, Australia

Andrew E. Grulich

Affiliation Kirby Institute, University of New South Wales, Sydney, Australia

Suzanne M. Garland

Affiliation Dept of Microbiology and Infectious Diseases, Royal Women’s Hospital, Department of Obstetrics and Gynaecology, University of Melbourne, Murdoch Children’s Research Institute, Melbourne, Australia

Catriona S. Bradshaw

Affiliations Melbourne Sexual Health Centre, Alfred Health, Melbourne, Australia, Dept of Epidemiology and Preventive Medicine, Monash University, Melbourne, Australia

Marcus Y. Chen

Affiliations Melbourne Sexual Health Centre, Alfred Health, Melbourne, Australia, School of Population Health, University of Melbourne, Melbourne, Australia

Christopher K. Fairley

Affiliations Melbourne Sexual Health Centre, Alfred Health, Melbourne, Australia, School of Population Health, University of Melbourne, Melbourne, Australia

Competing Interests

CKF, JSH and AEG have received honoraria from CSL Biotherapies. CKF and AEG have received research funding from CSL Biotherapies. CKF owns shares in CSL Biotherapies, the manufacturer of Gardasil, and has received honoraria from Merck. SMG has received advisory board fees and grant support from Commonwealth Serum Laboratories and GlaxoSmithKline, and lecture fees from Merck and Sanofi Pasteur MSD. In addition, she has received funding through her institutions to conduct HPV vaccine studies for GlaxoSmithKline and for Merck. AEG has received honoraria and travel funding from Merck, and sits on the Australian advisory board for the Gardasil vaccine. TRHR is a site investigator on a Merck-sponsored study. MYC, SNT, MJM and CSB, have no conflicts of interest. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CKF TRHR SNT MJM SMG MYC CSB JSH. Performed the experiments: LAV SNT TRHR MJM. Analyzed the data: TRHR JSH CKF AEG. Contributed reagents/materials/analysis tools: SNT SMG. Wrote the paper: TRHR CKF JSH AEG. Critical revision of manuscript: JSH LAV SNT MJM AEG CSB MYC SMG CKF.