About the Authors

Tsuyoshi Hamano

thamano@med.shimane-u.ac.jp

Affiliations Center for Community-Based Health Research and Education (COHRE), Organization for the Promotion of Project Research, Shimane University, Izumo, Japan, Department of Environmental and Preventive Medicine, Shimane University School of Medicine, Izumo, Japan

Yoshinari Kimura

Affiliation Department of Geography, Graduate School of Literature and Human Sciences, Osaka City University, Osaka, Japan

Miwako Takeda

Affiliations Center for Community-Based Health Research and Education (COHRE), Organization for the Promotion of Project Research, Shimane University, Izumo, Japan, Department of Environmental and Preventive Medicine, Shimane University School of Medicine, Izumo, Japan

Masayuki Yamasaki

Affiliations Center for Community-Based Health Research and Education (COHRE), Organization for the Promotion of Project Research, Shimane University, Izumo, Japan, Department of Environmental and Preventive Medicine, Shimane University School of Medicine, Izumo, Japan

Minoru Isomura

Affiliations Center for Community-Based Health Research and Education (COHRE), Organization for the Promotion of Project Research, Shimane University, Izumo, Japan, Department of Functional Pathology, Shimane University School of Medicine, Izumo, Japan

Toru Nabika

Affiliations Center for Community-Based Health Research and Education (COHRE), Organization for the Promotion of Project Research, Shimane University, Izumo, Japan, Department of Functional Pathology, Shimane University School of Medicine, Izumo, Japan

Kuninori Shiwaku

Affiliations Center for Community-Based Health Research and Education (COHRE), Organization for the Promotion of Project Research, Shimane University, Izumo, Japan, Department of Environmental and Preventive Medicine, Shimane University School of Medicine, Izumo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TH TN KS. Analyzed the data: TH YK. Contributed reagents/materials/analysis tools: TH MT MY MI. Wrote the paper: TH KS.