About the Authors

Erika Kague

Contributed equally to this work with: Erika Kague, Michael Gallagher

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Michael Gallagher

Contributed equally to this work with: Erika Kague, Michael Gallagher

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sally Burke

Affiliation Biology Department, Mount Saint Vincent University, Halifax, Nova Scotia, Canada

Michael Parsons

Affiliation McCusick–Nathans Institute of Genetic Medicine and Department of Surgery, The Johns Hopkins University, Baltimore, Maryland, United States of America

Tamara Franz-Odendaal

Affiliation Biology Department, Mount Saint Vincent University, Halifax, Nova Scotia, Canada

Shannon Fisher

sfisher4@upenn.edu

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SF MP. Performed the experiments: EK MG SB MP TFO SF. Analyzed the data: SF TFO. Contributed reagents/materials/analysis tools: MP MG. Wrote the paper: SF TFO.