About the Authors

Takashi Ishida

Affiliations RIKEN Plant Science Center, Tsurumi, Yokohama, Kanagawa, Japan, Graduate School of Biological Sciences, Nara Institute of Science and Technology, Ikoma, Nara, Japan

Mika Yoshimura

Affiliation RIKEN Plant Science Center, Tsurumi, Yokohama, Kanagawa, Japan

Kenji Miura

Affiliation Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Japan

Keiko Sugimoto

sugimoto@psc.riken.jp

Affiliation RIKEN Plant Science Center, Tsurumi, Yokohama, Kanagawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TI KM KS. Performed the experiments: TI MY KM. Wrote the paper: TI KM KS.