About the Authors

Shaoyuan Wu

shaoyuan5@gmail.com (SW); jmeng@amnh.org (JM)

Affiliation Department of Organismic and Evolutionary Biology and Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, United States of America

Wenyu Wu

Affiliation Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China

Fuchun Zhang

Affiliation Xinjiang Key Laboratory of Biological Resources and Genetic Engineering, Xinjiang University, Urumqi, Xinjiang, China

Jie Ye

Affiliation Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China

Xijun Ni

Affiliation Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing, China

Jimin Sun

Affiliation Key Lab of Cenozoic Geology and Environment, Institute of Geology and Geophysics, Chinese Academy of Sciences, Beijing, China

Scott V. Edwards

Affiliation Department of Organismic and Evolutionary Biology and Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, United States of America

Jin Meng

shaoyuan5@gmail.com (SW); jmeng@amnh.org (JM)

Affiliation Division of Paleontology, American Museum of Natural History, New York, New York, United States of America

Chris L. Organ

Affiliation Department of Organismic and Evolutionary Biology and Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SW SE JM. Performed the experiments: SW. Analyzed the data: SW JM CO. Contributed reagents/materials/analysis tools: WW FZ JY XN JS. Wrote the paper: SW CO.