About the Authors

Yongqiang Zhang

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, People's Republic of China

Xinwu Pei

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, People's Republic of China

Chao Zhang

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, People's Republic of China

Zifeng Lu

Affiliation State Key Laboratory of Grassland Agro-ecosystems, College of Pastoral Agriculture Science and Technology, Lanzhou University, Lanzhou, People's Republic of China

Zhixing Wang

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, People's Republic of China

Shirong Jia

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, People's Republic of China

Weimin Li

liwm518@hotmail.com

Affiliation Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YZ XP ZW SJ WL. Performed the experiments: YZ CZ ZL WL. Analyzed the data: XP ZW SJ WL. Contributed reagents/materials/analysis tools: YZ XP WL. Wrote the paper: YZ WL.