About the Authors

Andrew M. Ramey

aramey@usgs.gov

Affiliation U.S. Geological Survey, Alaska Science Center, Anchorage, Alaska, United States of America

Craig R. Ely

Affiliation U.S. Geological Survey, Alaska Science Center, Anchorage, Alaska, United States of America

Joel A. Schmutz

Affiliation U.S. Geological Survey, Alaska Science Center, Anchorage, Alaska, United States of America

John M. Pearce

Affiliation U.S. Geological Survey, Alaska Science Center, Anchorage, Alaska, United States of America

Darryl J. Heard

Affiliation College of Veterinary Medicine, University of Florida, Gainesville, Florida, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AMR CRE JAS JMP. Performed the experiments: CRE DJH. Analyzed the data: AMR CRE JAS. Wrote the paper: AMR CRE JAS JMP DJH.