About the Authors

Roberto A. Saenz

Current address: Institute of Integrative Biology, ETH Zürich, Switzerland

Affiliation Department of Applied Mathematics and Theoretical Physics, University of Cambridge, Cambridge, United Kingdom

Steve C. Essen

Affiliation Animal Health and Veterinary Laboratories Agency, United Kingdom; European Union/World Organisation for Animal Health/Food and Agriculture Organization Reference Laboratory for Avian Influenza and Newcastle Disease, Addlestone, Surrey, United Kingdom

Sharon M. Brookes

Affiliation Animal Health and Veterinary Laboratories Agency, United Kingdom; European Union/World Organisation for Animal Health/Food and Agriculture Organization Reference Laboratory for Avian Influenza and Newcastle Disease, Addlestone, Surrey, United Kingdom

Munir Iqbal

Affiliation Institute for Animal Health, Compton Laboratory, Compton, Newbury, Berkshire, United Kingdom

James L. N. Wood

Affiliation Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Bryan T. Grenfell

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

John W. McCauley

Affiliation Division of Virology, MRC National Institute for Medical Research, The Ridgeway, Mill Hill, London, United Kindom

Ian H. Brown

Affiliation Animal Health and Veterinary Laboratories Agency, United Kingdom; European Union/World Organisation for Animal Health/Food and Agriculture Organization Reference Laboratory for Avian Influenza and Newcastle Disease, Addlestone, Surrey, United Kingdom

Julia R. Gog

jrg20@cam.ac.uk

Affiliation Department of Applied Mathematics and Theoretical Physics, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BTG IHB JWM. Performed the experiments: MI SCE. Analyzed the data: IHB JRG JWM MI RAS SMB SCE. Wrote the paper: BTG JLNW JRG JWM RAS SMB.