About the Authors

Zhiling Guo

Affiliations Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Science, Guangzhou, Guangdong, China, Department of Marine Sciences, University of Connecticut, Groton, Connecticut, United States of America

Sheng Liu

Affiliation Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Science, Guangzhou, Guangdong, China

Simin Hu

Affiliation Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Science, Guangzhou, Guangdong, China

Tao Li

Affiliation Key Laboratory of Marine Bio-resources Sustainable Utilization, South China Sea Institute of Oceanology, Chinese Academy of Science, Guangzhou, Guangdong, China

Yousong Huang

Affiliation Department of Environmental Science, Ocean University of China, Qingdao, Shandong, China

Guangxing Liu

Affiliation Department of Environmental Science, Ocean University of China, Qingdao, Shandong, China

Huan Zhang

huan.zhang@uconn.edu (HZ); senjie.lin@uconn.edu (SL)

Affiliations Department of Marine Sciences, University of Connecticut, Groton, Connecticut, United States of America, Department of Environmental Science, Ocean University of China, Qingdao, Shandong, China

Senjie Lin

huan.zhang@uconn.edu (HZ); senjie.lin@uconn.edu (SL)

Affiliations Department of Marine Sciences, University of Connecticut, Groton, Connecticut, United States of America, Marine Biodiversity and Global Change Laboratory, Xiamen University, Xiamen, Fujian, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJL HZ. Performed the experiments: ZLG HZ SMH TL YSH. Analyzed the data: ZLG HZ SJL. Contributed reagents/materials/analysis tools: SJL SL GXL. Wrote the paper: ZLG HZ SJL.