About the Authors

Simon C. Watkins

Affiliation Department of Cell Biology, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Salony Maniar

Affiliation Department of Environmental and Occupational Health, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Mackenzie Mosher

Affiliation Department of Environmental and Occupational Health, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Beth L. Roman

Affiliation Department of Biological Sciences, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Michael Tsang

Affiliation Department of Developmental Biology, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Claudette M. St Croix

cls13@pitt.edu

Affiliations Department of Cell Biology, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Environmental and Occupational Health, The University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SCW CMS. Performed the experiments: SCW SM MM BLR MT CMS. Analyzed the data: SCW SM MM CMS. Contributed reagents/materials/analysis tools: SCW BLR MT CMS. Wrote the paper: SCW CMS. Critical editing of manuscript: BLR MT.