About the Authors

Roman J. Krawetz

rkrawetz@ucalgary.ca

Affiliations Department of Surgery, University of Calgary, Calgary, Alberta, Canada, Department of Cell Biology and Anatomy, University of Calgary, Calgary, Alberta, Canada

Yiru Elizabeth Wu

Affiliation Department of Surgery, University of Calgary, Calgary, Alberta, Canada

Liam Martin

Affiliation Department of Medicine, University of Calgary, Calgary, Alberta, Canada

Jerome B. Rattner

Affiliation Department of Cell Biology and Anatomy, University of Calgary, Calgary, Alberta, Canada

John R. Matyas

Affiliation Department of Comparative Biology and Experimental Medicine, Faculty of Veterinary Medicine, University of Calgary, Calgary, Alberta, Canada

David A. Hart

Affiliation Department of Surgery, University of Calgary, Calgary, Alberta, Canada

Competing Interests

The authors have the following interest: This study was partly funded by Pfizer Inc. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: RJK JBR JRM DAH. Performed the experiments: RJK YEW JBR JRM. Analyzed the data: RJK. Contributed reagents/materials/analysis tools: LM JRM. Wrote the paper: RJK YEW.