About the Authors

Nathan V. Whelan

nwhelan@crimson.ua.edu

Affiliation Department of Biological Sciences, University of Alabama, Tuscaloosa, Alabama, United States of America

Paul D. Johnson

Affiliation Alabama Aquatic Biodiversity Center, Alabama Department of Conservation and Natural Resources, Marion, Alabama, United States of America

Phil M. Harris

Affiliation Department of Biological Sciences, University of Alabama, Tuscaloosa, Alabama, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NW PJ PH. Performed the experiments: NW PJ. Analyzed the data: NW. Contributed reagents/materials/analysis tools: PJ PH. Wrote the paper: NW.