About the Authors

Timothy J. Ciarlo

tjc273@psu.edu

Affiliation Department of Entomology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Christopher A. Mullin

Affiliation Department of Entomology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

James L. Frazier

Affiliation Department of Entomology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Daniel R. Schmehl

Affiliation Department of Entomology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TJC CAM JLF DRS. Performed the experiments: TJC. Analyzed the data: TJC CAM JLF. Contributed reagents/materials/analysis tools: TJC CAM JLF DRS. Wrote the paper: TJC. Developed automated PER protocol: DRS JLF. Reviewed manuscript: CAM JLF DRS.