About the Authors

Hiroko Eda-Fujiwara

Contributed equally to this work with: Hiroko Eda-Fujiwara, Takuya Imagawa

hirofujiwa@fc.jwu.ac.jp; sbhtake@ipc.shizuoka.ac.jp

Affiliations Department of Chemical & Biological Sciences, Japan Women’s University, Bunkyo-ku, Tokyo, Japan, Japan Society for the Promotion of Science, Chiyoda-ku, Tokyo, Japan

Takuya Imagawa

Contributed equally to this work with: Hiroko Eda-Fujiwara, Takuya Imagawa

Affiliation Department of Biology, Faculty of Science, Shizuoka University, Shizuoka, Japan

Masanori Matsushita

Affiliation Department of Biology, Faculty of Science, Shizuoka University, Shizuoka, Japan

Yasushi Matsuda

Affiliation Department of Biology, Faculty of Science, Shizuoka University, Shizuoka, Japan

Hiro-Aki Takeuchi

hirofujiwa@fc.jwu.ac.jp; sbhtake@ipc.shizuoka.ac.jp

Affiliation Department of Biology, Faculty of Science, Shizuoka University, Shizuoka, Japan

Ryohei Satoh

Affiliation Department of Physiology, Kitasato University School of Medicine, Kanagawa, Japan

Aiko Watanabe

Affiliation Department of Chemical & Biological Sciences, Japan Women’s University, Bunkyo-ku, Tokyo, Japan

Matthijs A. Zandbergen

Affiliation Behavioural Biology and Helmholtz Institute, Utrecht University, Utrecht, The Netherlands

Kazuchika Manabe

Affiliation Graduate School of Social and Cultural Studies, Nihon University, Saitama, Japan

Takashi Kawashima

Affiliation Graduate School of Social and Cultural Studies, Nihon University, Saitama, Japan

Johan J. Bolhuis

Affiliation Behavioural Biology and Helmholtz Institute, Utrecht University, Utrecht, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HE-F TI H-AT JJB. Performed the experiments: HE-F TI MM YM RS AW. Analyzed the data: RS MAZ. Contributed reagents/materials/analysis tools: KM TK. Wrote the paper: HE-F RS JJB.