About the Authors

Marcos G. Frank

mgf@mail.med.upenn.edu (MGF); Jean.Boal@millersville.edu (JGB)

Affiliation Department of Neuroscience, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Robert H. Waldrop

Affiliation Department of Biology, Millersville University, Lancaster, Pennsylvania, United States of America

Michelle Dumoulin

Affiliation Department of Neuroscience, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sara Aton

Affiliation Department of Neuroscience, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jean G. Boal

mgf@mail.med.upenn.edu (MGF); Jean.Boal@millersville.edu (JGB)

Affiliation Department of Biology, Millersville University, Lancaster, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MGF JGB. Performed the experiments: RW MD SA. Analyzed the data: RW MGF JGB. Contributed reagents/materials/analysis tools: MGF JGB. Wrote the paper: MGF JGB.