About the Authors

Masahiko Arikata

Affiliations Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan, Department of Otorhinolaryngology-Head and Neck Surgery, Shiga University of Medical Science, Otsu, Japan

Yasushi Itoh

yasushii@belle.shiga-med.ac.jp

Affiliation Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan

Masatoshi Okamatsu

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Toshinaga Maeda

Affiliation Division of Molecular Medical Biochemistry, Shiga University of Medical Science, Otsu, Japan

Takashi Shiina

Affiliation Department of Molecular Life Science, Division of Basic Medical Science and Molecular Medicine, Tokai University School of Medicine, Isehara, Japan

Keiko Tanaka

Affiliation Department of Molecular Life Science, Division of Basic Medical Science and Molecular Medicine, Tokai University School of Medicine, Isehara, Japan

Shingo Suzuki

Affiliation Department of Molecular Life Science, Division of Basic Medical Science and Molecular Medicine, Tokai University School of Medicine, Isehara, Japan

Misako Nakayama

Affiliation Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan

Yoshihiro Sakoda

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Hirohito Ishigaki

Affiliation Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan

Ayato Takada

Affiliation Research Center for Zoonosis Control, Hokkaido University, Sapporo, Japan

Hideaki Ishida

Affiliation Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan

Kosuke Soda

Affiliation Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan

Van Loi Pham

Affiliation Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan

Hideaki Tsuchiya

Affiliation Research Center for Animal Life Science, Shiga University of Medical Science, Otsu, Japan

Shinichiro Nakamura

Affiliation Research Center for Animal Life Science, Shiga University of Medical Science, Otsu, Japan

Ryuzo Torii

Affiliation Research Center for Animal Life Science, Shiga University of Medical Science, Otsu, Japan

Takeshi Shimizu

Affiliation Department of Otorhinolaryngology-Head and Neck Surgery, Shiga University of Medical Science, Otsu, Japan

Hidetoshi Inoko

Affiliation Department of Molecular Life Science, Division of Basic Medical Science and Molecular Medicine, Tokai University School of Medicine, Isehara, Japan

Iwao Ohkubo

Affiliation Division of Molecular Medical Biochemistry, Shiga University of Medical Science, Otsu, Japan

Hiroshi Kida

Affiliations Laboratory of Microbiology, Department of Disease Control, Graduate School of Veterinary Medicine, Hokkaido University, Sapporo, Japan, Research Center for Zoonosis Control, Hokkaido University, Sapporo, Japan

Kazumasa Ogasawara

Affiliation Division of Pathology and Disease Regulation, Department of Pathology, Shiga University of Medical Science, Otsu, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YI YS T. Shiina HK KO. Performed the experiments: MA YI MO TM T. Shiina KT SS MN H. Ishigaki H. Ishida KS VLP. Analyzed the data: MA YI MO TM T. Shiina H. Inoko. Contributed reagents/materials/analysis tools: AT HT SN RT T. Shimizu IO. Wrote the paper: YI KO.