About the Authors

Kazutaka Ohi

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Core Research for Evolutionary Science and Technology (CREST) of Japan Science and Technology Agency (JST), Saitama, Japan, National Hospital Organization, Yamato Mental-Medical Center, Nara, Japan

Ryota Hashimoto

hashimor@psy.med.osaka-u.ac.jp

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Core Research for Evolutionary Science and Technology (CREST) of Japan Science and Technology Agency (JST), Saitama, Japan, Molecular Research Center for Children's Mental Development, United Graduate School of Child Development, Osaka University, Kanazawa University and Hamamatsu University School of Medicine, Osaka, Japan

Takanobu Nakazawa

Affiliations Division of Oncology, Institute of Medical Science, University of Tokyo, Tokyo, Japan, Department of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Takeya Okada

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Core Research for Evolutionary Science and Technology (CREST) of Japan Science and Technology Agency (JST), Saitama, Japan

Yuka Yasuda

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Core Research for Evolutionary Science and Technology (CREST) of Japan Science and Technology Agency (JST), Saitama, Japan

Hidenaga Yamamori

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Core Research for Evolutionary Science and Technology (CREST) of Japan Science and Technology Agency (JST), Saitama, Japan, Department of Molecular Neuropsychiatry, Osaka University Graduate School of Medicine, Osaka, Japan

Motoyuki Fukumoto

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Core Research for Evolutionary Science and Technology (CREST) of Japan Science and Technology Agency (JST), Saitama, Japan

Satomi Umeda-Yano

Affiliation Department of Molecular Neuropsychiatry, Osaka University Graduate School of Medicine, Osaka, Japan

Masao Iwase

Affiliation Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan

Hiroaki Kazui

Affiliation Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan

Tadashi Yamamoto

Affiliation Division of Oncology, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Masanobu Kano

Affiliation Department of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Masatoshi Takeda

Affiliations Department of Psychiatry, Osaka University Graduate School of Medicine, Osaka, Japan, Molecular Research Center for Children's Mental Development, United Graduate School of Child Development, Osaka University, Kanazawa University and Hamamatsu University School of Medicine, Osaka, Japan

Competing Interests

Ryota Hashimoto is an academic editor of this journal. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KO RH TN TY MK MT. Performed the experiments: TO YY HY MF SU. Analyzed the data: KO RH TN MI HK. Contributed reagents/materials/analysis tools: TO YY HY MF SU MI HK TY MK MT. Wrote the paper: KO RH TN.