About the Authors

Desmond D. Campbell

These authors are joint first authors on this work.

Affiliation Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Maria V. Parra

These authors are joint first authors on this work.

Affiliations Department of Genetics, Evolution and Environment, University College London, London, United Kingdom, Laboratorio de Genética Molecular, Universidad de Antioquía, Medellín, Colombia

Constanza Duque

These authors are joint first authors on this work.

Affiliations Department of Genetics, Evolution and Environment, University College London, London, United Kingdom, Laboratorio de Genética Molecular, Universidad de Antioquía, Medellín, Colombia

Natalia Gallego

Affiliation Laboratorio de Genética Molecular, Universidad de Antioquía, Medellín, Colombia

Liliana Franco

Affiliation Laboratorio de Genética Molecular, Universidad de Antioquía, Medellín, Colombia

Arti Tandon

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Tábita Hünemeier

Affiliation Departamento de Genética, Universidad Federal do Rio Grande do Sul, Porto Alegre, Brazil

Cátira Bortolini

Affiliation Departamento de Genética, Universidad Federal do Rio Grande do Sul, Porto Alegre, Brazil

Alberto Villegas

Affiliation Laboratorio de Genética Molecular, Universidad de Antioquía, Medellín, Colombia

Gabriel Bedoya

Affiliation Laboratorio de Genética Molecular, Universidad de Antioquía, Medellín, Colombia

Mark I. McCarthy

Affiliations Oxford Centre for Diabetes, Endocrinology and Metabolism, University of Oxford, Churchill Hospital, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Oxford NIHR Biomedical Research Centre, Churchill Hospital, Oxford, United Kingdom

Alkes Price

Affiliation Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America

David Reich

Affiliations Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America, Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Andrés Ruiz-Linares

a.ruizlin@ucl.ac.uk

Affiliation Department of Genetics, Evolution and Environment, University College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DDC CB AV GB MM AP DR AR-L. Performed the experiments: MVP CD NG LF TH. Analyzed the data: DDC MVP CD AT DR AR-L. Wrote the paper: DDC AR-L.