About the Authors

William R. Swindell

wswindell@genetics.med.harvard.edu

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Andrew Johnston

Affiliation Department of Dermatology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Liou Sun

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Xianying Xing

Affiliation Department of Dermatology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Gary J. Fisher

Affiliation Department of Dermatology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Martha L. Bulyk

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America, Department of Pathology, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America, Division of Health Sciences and Technology, Harvard Medical School, Boston, Massachusetts, United States of America

James T. Elder

Affiliation Department of Dermatology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Johann E. Gudjonsson

Affiliation Department of Dermatology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WRS AJ GJF JTE JEG. Performed the experiments: WRS AJ LS XX JEG. Analyzed the data: WRS AJ MLB JEG. Contributed reagents/materials/analysis tools: LS. Wrote the paper: WRS AJ MLB JTE JEG.