About the Authors

Ali H. Ziyab

Affiliations Department of Epidemiology and Biostatistics, Norman J. Arnold School of Public Health, University of South Carolina, Columbia, South Carolina, United States of America, Department of Community Medicine and Behavioral Sciences, Faculty of Medicine, Kuwait University, Kuwait City, Kuwait

Wilfried Karmaus

karmaus@sc.edu

Affiliation Department of Epidemiology and Biostatistics, Norman J. Arnold School of Public Health, University of South Carolina, Columbia, South Carolina, United States of America

Mitra Yousefi

Affiliation Department of Epidemiology and Biostatistics, Norman J. Arnold School of Public Health, University of South Carolina, Columbia, South Carolina, United States of America

Susan Ewart

Affiliation College of Veterinary Medicine, Michigan State University, East Lansing, Michigan, United States of America

Eric Schauberger

Affiliations Genetics Graduate Program, Michigan State University, East Lansing, Michigan, United States of America, College of Osteopathic Medicine, Michigan State University, East Lansing, Michigan, United States of America

John W. Holloway

Affiliations Academic Units of Clinical and Experimental Medicine, Faculty of Medicine, University of Southampton, Southampton, United Kingdom, Academic Units of Human Genetics, Faculty of Medicine, University of Southampton, Southampton, United Kingdom

Hongmei Zhang

Affiliation Department of Epidemiology and Biostatistics, Norman J. Arnold School of Public Health, University of South Carolina, Columbia, South Carolina, United States of America

Syed Hasan Arshad

Affiliations Academic Units of Clinical and Experimental Medicine, Faculty of Medicine, University of Southampton, Southampton, United Kingdom, David Hide Asthma and Allergy Research Centre, Isle of Wight, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: AHZ WK MY HZ. Wrote the paper: AHZ WK MY. Conceived and designed the epidemiological study: AHZ SHA WK ES Performed the examinations and measurements: AHZ SE ES. Revised the manuscript: HZ SE JWH WK.