About the Authors

Yusuke Hama

Contributed equally to this work with: Yusuke Hama, Tokuhiro Chano

Affiliation Department of Clinical Laboratory Medicine, Shiga University of Medical Science, Shiga, Japan

Tokuhiro Chano

Contributed equally to this work with: Yusuke Hama, Tokuhiro Chano

chano@belle.shiga-med.ac.jp

Affiliation Department of Clinical Laboratory Medicine, Shiga University of Medical Science, Shiga, Japan

Takuma Inui

Affiliation Department of Clinical Laboratory Medicine, Shiga University of Medical Science, Shiga, Japan

Kyoichi Matsumoto

Affiliation Mikuri Immunology Laboratory, Osaka, Japan

Hidetoshi Okabe

Affiliation Department of Clinical Laboratory Medicine, Shiga University of Medical Science, Shiga, Japan

Competing Interests

The authors have the following competing interest. Kyoichi Matsumoto is an employee of Mikuri Immunology Laboratory, Inc. and collaborated with the other authors in immunizing mice and preparing hybridomas for the monoclonal antibodies. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: TC HO. Performed the experiments: YH TC TI KM HO. Analyzed the data: YH TC. Contributed reagents/materials/analysis tools: KM. Wrote the paper: YH TC.