About the Authors

Steven B. Roberts

sr320@uw.edu

Affiliation School of Aquatic and Fishery Sciences, University of Washington, Seattle, Washington, United States of America

Lorenz Hauser
Lisa W. Seeb
James E. Seeb

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SBR LH LWS JES. Performed the experiments: SBR. Analyzed the data: SBR LH LWS. Contributed reagents/materials/analysis tools: SBR JES. Wrote the paper: SBR LH LWS JES.