About the Authors

Javier Simón-Sánchez

Contributed equally to this work with: Javier Simón-Sánchez, Laura L. Kilarski

Affiliation Section of Medical Genomics, Department of Clinical Genetics, VU University Medical Centre, Amsterdam, The Netherlands

Laura L. Kilarski

Contributed equally to this work with: Javier Simón-Sánchez, Laura L. Kilarski

Affiliations MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom, Department of Psychological Medicine & Neurology, Cardiff University School of Medicine, Cardiff, United Kingdom

Michael A. Nalls

Affiliation Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

Maria Martinez

Affiliations Inserm, UMR 1043, Toulouse, France, Paul Sabatier University, Toulouse, France

Claudia Schulte

Affiliation Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, and DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Peter Holmans

Affiliations MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom, Department of Psychological Medicine & Neurology, Cardiff University School of Medicine, Cardiff, United Kingdom

International Parkinson's Disease Genomics Consortium

a Membership of the International Parkinson's Disease Genomics Consortium is provided in the Acknowledgments.

Wellcome Trust Case Control Consortium

b Membership of the Wellcome Trust Case Control Consortium is provided in supporting information.

Affiliation MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom

Thomas Gasser

Affiliation Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, and DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

John Hardy

Affiliation Department of Molecular Neuroscience, UCL Institute of Neurology, London, United Kingdom

Andrew B. Singleton

Affiliation Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

Nicholas W. Wood

Affiliation Department of Molecular Neuroscience, UCL Institute of Neurology, London, United Kingdom

Alexis Brice

Affiliations Université Pierre et Marie Curie-Paris6, Centre de Recherche de l'Institut du Cerveau et de la Moelle épinière, UMR-S975, Paris, France, Inserm, U975, Paris, France, Cnrs, UMR 7225, Paris, France

Peter Heutink

Affiliation Section of Medical Genomics, Department of Clinical Genetics, VU University Medical Centre, Amsterdam, The Netherlands

Nigel Williams

williamsnm@cf.ac.uk (NW); morrishr@cf.ac.uk (HRM)

Affiliations MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom, Department of Psychological Medicine & Neurology, Cardiff University School of Medicine, Cardiff, United Kingdom

Huw R. Morris

williamsnm@cf.ac.uk (NW); morrishr@cf.ac.uk (HRM)

Affiliations MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom, Department of Psychological Medicine & Neurology, Cardiff University School of Medicine, Cardiff, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HRM NMW LLK JS-S CS. Performed the experiments: JS-S LLK. Analyzed the data: LLK JS-S. Contributed reagents/materials/analysis tools: CS. Wrote the paper: LLK JS-S HRM NMW. Contributed samples: TG JH ABS NWW AB PH NMW HRM WTCCC2. Critically reviewed manuscript: MAN MM PAH JH AB NMW HRM JS-S LLK.