About the Authors

Byoung-Shik Shim

Affiliations Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea, Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Sciences, Center for Agricultural Biomaterials, Seoul National University, Seoul, Republic of Korea

Young Ki Choi

Affiliation College of Medicine and Medical Research Institute, Chungbuk National University, Cheongju, Republic of Korea

Cheol-Heui Yun

Affiliation Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Sciences, Center for Agricultural Biomaterials, Seoul National University, Seoul, Republic of Korea

Eu-Gene Lee

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Yoon Seong Jeon

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Sung-Moo Park

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

In Su Cheon

Affiliations Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea, Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Sciences, Center for Agricultural Biomaterials, Seoul National University, Seoul, Republic of Korea

Dong-Hyun Joo

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Chung Hwan Cho

Affiliations Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea, Department of Agricultural Biotechnology, Research Institute for Agriculture and Life Sciences, Center for Agricultural Biomaterials, Seoul National University, Seoul, Republic of Korea

Min-Suk Song

Affiliation College of Medicine and Medical Research Institute, Chungbuk National University, Cheongju, Republic of Korea

Sang-Uk Seo

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Young-Ho Byun

Affiliation Department of Biotechnology, College of Engineering, Yonsei University, Seoul, Republic of Korea

Hae-Jung Park

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Haryoung Poo

Affiliation Korean Research Institute of Bioscience and Biotechnology, Daejeon, Republic of Korea

Baik Lin Seong

Affiliation Department of Biotechnology, College of Engineering, Yonsei University, Seoul, Republic of Korea

Jae Ouk Kim

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Huan Huu Nguyen

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Konrad Stadler

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Dong Wook Kim

Affiliation Department of Pharmacy, College of Pharmacy, Hanyang University, Kyeonggi-do, Republic of Korea

Kee-Jong Hong

Affiliation Division of Influenza Viruses, Korea National Institute of Health, Osong, Republic of Korea

Cecil Czerkinsky

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Man Ki Song

mksong@ivi.int

Affiliation Laboratory Science Division, International Vaccine Institute, Seoul, Republic of Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MKS CC. Performed the experiments: BSS EGL SMP HJP ISC MSS DHJ SUS YHB. Analyzed the data: HHN DWK. Contributed reagents/materials/analysis tools: YSJ HHN HRP BLS YKC CC. Wrote the paper: BSS YKC CHY CHC KS MKS. Advised editing and discussion of the manuscript: MKS JOK KJH KS HHN CHY.