About the Authors

Jennifer A. Schumacher

Contributed equally to this work with: Jennifer A. Schumacher, Megumi Hashiguchi, Vu H. Nguyen

Current address: Division of Developmental Biology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Megumi Hashiguchi

Contributed equally to this work with: Jennifer A. Schumacher, Megumi Hashiguchi, Vu H. Nguyen

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Vu H. Nguyen

Contributed equally to this work with: Jennifer A. Schumacher, Megumi Hashiguchi, Vu H. Nguyen

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Mary C. Mullins

mullins@mail.med.upenn.edu

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JAS MH VHN MCM. Performed the experiments: JAS MH VHN. Analyzed the data: JAS MH VHN MCM. Contributed reagents/materials/analysis tools: JAS MH VHN MCM. Wrote the paper: JAS MH VHN MCM.