About the Authors

Meredith G. Schafer

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Andrew A. Ross

Affiliation Department of Biological Sciences, North Dakota State University, Fargo, North Dakota, United States of America

Jason P. Londo

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Connie A. Burdick

Affiliation Western Ecology Division, National Health and Environmental Effects Research Laboratory, U.S. Environmental Protection Agency, Corvallis, Oregon, United States of America

E. Henry Lee

Affiliation Western Ecology Division, National Health and Environmental Effects Research Laboratory, U.S. Environmental Protection Agency, Corvallis, Oregon, United States of America

Steven E. Travers

Affiliation Department of Biological Sciences, North Dakota State University, Fargo, North Dakota, United States of America

Peter K. Van de Water

Affiliation Earth and Environmental Sciences Department, California State University, Fresno, California, United States of America

Cynthia L. Sagers

csagers@uark.edu

Affiliation Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MGS AAR JPL EHL CLS. Performed the experiments: MGS AAR JPL SET CLS. Analyzed the data: MGS CAB EHL JPL CLS. Contributed reagents/materials/analysis tools: EHL PKV CLS. Wrote the paper: MGS JPL CAB EHL CLS. Publication costs: PKV CLS.