About the Authors

Preston Foerder

Affiliation Biopsychology and Behavioral Neuroscience Program, The Graduate Center, The City University of New York, New York, New York, United States of America

Marie Galloway

Affiliation Smithsonian National Zoological Park, Washington, D.C., United States of America

Tony Barthel

Affiliation Smithsonian National Zoological Park, Washington, D.C., United States of America

Donald E. Moore III

Affiliation Smithsonian National Zoological Park, Washington, D.C., United States of America

Diana Reiss

dlr28@columbia.edu

Affiliations Biopsychology and Behavioral Neuroscience Program, The Graduate Center, The City University of New York, New York, New York, United States of America, Department of Psychology, Hunter College, The City University of New York, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PF DR. Performed the experiments: PF MG TB. Analyzed the data: PF DR. Wrote the paper: PF DR. Animal management: MG TB DEM. Facilitated NZP research: DEM.