About the Authors

Yanchun Liang

Affiliation Key Laboratory of Symbol Computation and Knowledge Engineering of Ministry of Education, College of Computer Science and Technology, Jilin University, Changchun, China

Fan Zhang

Affiliation Key Laboratory of Symbol Computation and Knowledge Engineering of Ministry of Education, College of Computer Science and Technology, Jilin University, Changchun, China

Juexin Wang

Affiliation Key Laboratory of Symbol Computation and Knowledge Engineering of Ministry of Education, College of Computer Science and Technology, Jilin University, Changchun, China

Trupti Joshi

Affiliation Digital Biology Laboratory, Computer Science Department and Christopher S. Bond Life Sciences Center, University of Missouri, Columbia, Missouri, United States of America

Yan Wang

wy6868@hotmail.com (YW); xudong@missouri.edu (DX)

Affiliation Key Laboratory of Symbol Computation and Knowledge Engineering of Ministry of Education, College of Computer Science and Technology, Jilin University, Changchun, China

Dong Xu

wy6868@hotmail.com (YW); xudong@missouri.edu (DX)

Affiliations Key Laboratory of Symbol Computation and Knowledge Engineering of Ministry of Education, College of Computer Science and Technology, Jilin University, Changchun, China, Digital Biology Laboratory, Computer Science Department and Christopher S. Bond Life Sciences Center, University of Missouri, Columbia, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Performed the experiments: FZ JW. Analyzed the data: TJ YW. Wrote the paper: FZ YL DX. Conceived and designed the study: FZ YW YL. Guided the method development and data analysis: DX YW YL.