About the Authors

Denver W. Fowler

df9465@yahoo.co.uk

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Holly N. Woodward

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Elizabeth A. Freedman

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Peter L. Larson

Affiliations Black Hills Institute of Geological Research, Inc., Hill City, South Dakota, United States of America, School of Earth, Atmospheric and Environmental Sciences, University of Manchester, Manchester, United Kingdom

John R. Horner

Affiliation Museum of the Rockies and Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Competing Interests

We have the following competing interest: Peter L. Larson is an employee of Black Hills Institute of Geological Research Inc. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: DWF HNW EAF PLL. Performed the experiments: DWF HNW EAF PLL. Analyzed the data: DWF HNW EAF PLL JRH. Contributed reagents/materials/analysis tools: JRH. Wrote the paper: DWF HNW EAF PLL.