About the Authors

Erin L. Willis

ewillis@memphiszoo.org

Affiliation Department of Conservation and Research, Memphis Zoological Society, Memphis, Tennessee, United States of America

David C. Kersey

Current address: College of Veterinary Medicine, Western University of Health Sciences, Pomona, California, United States of America

Affiliation Department of Reproductive Sciences, Center for Species Survival, Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

Barbara S. Durrant

Affiliation Reproductive Physiology Division, Institute for Conservation Research, San Diego Zoo Global, Escondido, California, United States of America

Andrew J. Kouba

Affiliation Department of Conservation and Research, Memphis Zoological Society, Memphis, Tennessee, United States of America

Competing Interests

The Memphis Zoo has filed a provisional patent application associated with the applications of urinary ceruloplasmin as a non-invasive marker of pregnancy and pregnancy loss. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ELW. Performed the experiments: ELW. Analyzed the data: ELW. Contributed reagents/materials/analysis tools: DCK BSD AJK. Wrote the manuscript: ELW. Commented on the paper: DCK BSD AJK.