About the Authors

Megumi Kaneko

Affiliation Laboratory for Neural Cell Polarity, RIKEN Brain Science Institute, Wako, Saitama, Japan

Kazuhiko Yamaguchi

Affiliation Laboratory for Memory and Learning, RIKEN Brain Science Institute, Wako, Saitama, Japan

Mototsugu Eiraku

Affiliation Laboratory for Neural Cell Polarity, RIKEN Brain Science Institute, Wako, Saitama, Japan

Motohiko Sato

Affiliation Institute for Integrated Cell-Material Sciences, Kyoto University, Sakyo-ku, Kyoto, Japan

Norio Takata

Affiliation Hirase Research Unit, RIKEN Brain Science Institute, Wako, Saitama, Japan

Yoshimoto Kiyohara

Affiliation Laboratory for Neural Architecture, RIKEN Brain Science Institute, Wako, Saitama, Japan

Masayoshi Mishina

Affiliation Department of Molecular Neurobiology and Pharmacology, Graduate School of Medicine, University of Tokyo, Bunkyo-ku, Tokyo, Japan

Hajime Hirase

Affiliation Hirase Research Unit, RIKEN Brain Science Institute, Wako, Saitama, Japan

Tsutomu Hashikawa

Affiliation Laboratory for Neural Architecture, RIKEN Brain Science Institute, Wako, Saitama, Japan

Mineko Kengaku

kengaku@icems.kyoto-u.ac.jp

Current address: Institute for Integrated Cell-Material Sciences, Kyoto University, Yoshida-honmachi, Sakyo-ku, Kyoto, Japan

Affiliations Laboratory for Neural Cell Polarity, RIKEN Brain Science Institute, Wako, Saitama, Japan, Institute for Integrated Cell-Material Sciences, Kyoto University, Sakyo-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: M. Kengaku HH TH. Performed the experiments: M. Kaneko KY ME MS NT TH YK. Analyzed the data: M. Kaneko KY HH TH M. Kengaku. Contributed reagents/materials/analysis tools: MM. Wrote the paper: M. Kengaku.