About the Authors

Chia-Chyi Liu

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Meng-Shin Guo

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Fion Hsiao-Yu Lin

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Kuang-Nan Hsiao

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Kate Hsuen-Wen Chang

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Ai-Hsiang Chou

Affiliation Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Yu-Chao Wang

Affiliation Center for Nanomedicine Research, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Yu-Ching Chen

Affiliation Center for Nanomedicine Research, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Chung-Shi Yang

Affiliation Center for Nanomedicine Research, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan

Pele Choi-Sing Chong

pelechong@nhri.org.tw

Affiliations Vaccine Research and Development Center, National Health Research Institutes, Zhunan Town, Miaoli County, Taiwan, Graduate Institute of Immunology, China Medical University, Taichung, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: C-CL A-HC PC-SC. Performed the experiments: C-CL M-SG FH-YL K-NH KH-WC A-HC Y-CW Y-CC. Analyzed the data: C-CL A-HC C-SY PC-SC. Contributed reagents/materials/analysis tools: C-CL FH-YL A-HC CSY. Wrote the paper: C-CL A-HC PC-SC.