About the Authors

Tina L. Tootle

tina-tootle@uiowa.edu

Affiliation Department of Anatomy and Cell Biology, Roy J. and Lucille Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Dianne Williams

Affiliation Department of Embryology, Howard Hughes Medical Institute/Carnegie Institution, Baltimore, Maryland, United States of America

Alexander Hubb

Affiliation Department of Anatomy and Cell Biology, Roy J. and Lucille Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Rebecca Frederick

Affiliation Department of Embryology, Howard Hughes Medical Institute/Carnegie Institution, Baltimore, Maryland, United States of America

Allan Spradling

Affiliation Department of Embryology, Howard Hughes Medical Institute/Carnegie Institution, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TLT AS. Performed the experiments: TLT DW RF AH. Analyzed the data: TLT AS. Wrote the paper: TLT AS.