About the Authors

Patrick C. McHugh

Affiliation Department of Biology and Biochemistry, University of Bath, Claverton Down, Bath, United Kingdom

Josephine A. Wright

Affiliation Department of Biology and Biochemistry, University of Bath, Claverton Down, Bath, United Kingdom

David R. Brown

bssdrb@bath.ac.uk

Affiliation Department of Biology and Biochemistry, University of Bath, Claverton Down, Bath, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PM JW DB. Performed the experiments: PM. Contributed reagents/materials/analysis tools: PM DB. Wrote the paper: PM.