About the Authors

Michael H. H. Price

pricem@uvic.ca

Affiliations Department of Biology, University of Victoria, Victoria, Canada, Raincoast Conservation Foundation, Sidney, Canada

Stan L. Proboszcz

Affiliation Watershed Watch Salmon Society, Coquitlam, Canada

Rick D. Routledge

Affiliation Department of Statistics and Actuarial Science, Simon Fraser University, Burnaby, Canada

Allen S. Gottesfeld

Affiliation Skeena Fisheries Commission, Hazelton, Canada

Craig Orr

Affiliation Watershed Watch Salmon Society, Coquitlam, Canada

John D. Reynolds

Affiliation Earth to Ocean Research Group, Department of Biology, Simon Fraser University, Burnaby, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MP RR AG. Performed the experiments: MP AG. Analyzed the data: MP RR. Contributed reagents/materials/analysis tools: MP SP RR AG. Wrote the paper: MP SP RR AG CO JR.