About the Authors

Morgan J. Trimble

Affiliation Conservation Ecology Research Unit, Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa

Rudi J. van Aarde

rjvaarde@zoology.up.ac.za

Affiliation Conservation Ecology Research Unit, Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa

Competing Interests

The authors have been funded in part by a commercial source, Richards Bay Minerals. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MJT RJvA. Performed the experiments: MJT RJvA. Analyzed the data: MJT. Wrote the paper: MJT RJvA.