About the Authors

Marc Friedli

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School and University Hospitals of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Isabelle Barde

Affiliations School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne (EPFL), Lausanne, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Mélanie Arcangeli

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School and University Hospitals of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Sonia Verp

Affiliations School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne (EPFL), Lausanne, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Alexandra Quazzola

Affiliations School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne (EPFL), Lausanne, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Jozsef Zakany

Affiliations Department of Zoology and Animal Biology, University of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Nathalie Lin-Marq

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School and University Hospitals of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Daniel Robyr

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School and University Hospitals of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Catia Attanasio

Current address: Genomics Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School and University Hospitals of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

François Spitz

Current address: Developmental Biology Program, European Molecular Biology Laboratory, Heidelberg, Germany

Affiliations Department of Zoology and Animal Biology, University of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Denis Duboule

didier.trono@epfl.ch (DT); stylianos.antonarakis@unige.ch (SEA); denis.duboule@unige.ch (DD)

Affiliations School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne (EPFL), Lausanne, Switzerland, Department of Zoology and Animal Biology, University of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Didier Trono

didier.trono@epfl.ch (DT); stylianos.antonarakis@unige.ch (SEA); denis.duboule@unige.ch (DD)

Affiliations School of Life Sciences, Ecole Polytechnique Fédérale de Lausanne (EPFL), Lausanne, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Stylianos E. Antonarakis

didier.trono@epfl.ch (DT); stylianos.antonarakis@unige.ch (SEA); denis.duboule@unige.ch (DD)

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School and University Hospitals of Geneva, Geneva, Switzerland, “Frontiers in Genetics”, National Centre for Competence in Research, Bern, Switzerland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MF IB DR JZ FS DD DT SEA. Performed the experiments: MF MA SV AQ FS JZ IB NLM DR CA. Analyzed the data: MF FS IB DR DD DT SEA. Wrote the paper: MF FS DD DT SEA.