About the Authors

Joni Macdonald

Affiliation The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

James D. Glover

Affiliation The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

Lorna Taylor

Affiliation The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

Helen M. Sang

Affiliation The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

Michael J. McGrew

mike.mcgrew@roslin.ed.ac.uk

Affiliation The Roslin Institute and Royal Dick School of Veterinary Studies, University of Edinburgh, Roslin, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JM JDG HMS MJM. Performed the experiments: JM LT JDG MJM. Analyzed the data: JM JDG HMS LT MJM. Contributed reagents/materials/analysis tools: MJM. Wrote the paper: JM JDG HMS MJM.