About the Authors

Anne M. Wertheimer

Affiliation Arizona Center on Aging, University of Arizona, Tucson, Arizona, United States of America

Jennifer L. Uhrlaub

Affiliations Arizona Center on Aging, University of Arizona, Tucson, Arizona, United States of America, Department of Immunobiology, University of Arizona, Tucson, Arizona, United States of America

Alec Hirsch

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Guruprasad Medigeshi

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Jerald Sprague

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Alfred Legasse

Affiliation Oregon National Primate Research Center, Oregon Health and Science University, Beaverton, Oregon, United States of America

Jennifer Wilk

Affiliation Oregon National Primate Research Center, Oregon Health and Science University, Beaverton, Oregon, United States of America

Clayton A. Wiley

Affiliation University of Pittsburgh Medical Center, Pittsburgh, Pennsylvania, United States of America

Peter Didier

Affiliation Tulane National Primate Research Center, Covington, Louisiana, United States of America

Robert B. Tesh

Affiliation University of Texas Medical Branch, Galveston, Texas, United States of America

Kristy O. Murray

Affiliation University of Texas Health Science Center at Houston, School of Public Health, Houston, Texas, United States of America

Michael K. Axthelm

Affiliation Oregon National Primate Research Center, Oregon Health and Science University, Beaverton, Oregon, United States of America

Scott W. Wong

Affiliations Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America, Oregon National Primate Research Center, Oregon Health and Science University, Beaverton, Oregon, United States of America

Janko Nikolich-Žugich

nikolich@email.arizona.edu

Affiliations Arizona Center on Aging, University of Arizona, Tucson, Arizona, United States of America, Department of Immunobiology, University of Arizona, Tucson, Arizona, United States of America, Oregon National Primate Research Center, Oregon Health and Science University, Beaverton, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JNZ. Performed the experiments: AW JU AH GM JS. Analyzed the data: AW AH GM JS SW KM CW PD RT JNZ. Contributed reagents/materials/analysis tools: CW PD SW RT. Wrote the paper: AW JNZ. AL JW MA.