About the Authors

Timothy K. Williams

Contributed equally to this work with: Timothy K. Williams, Christina L. Costantino

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Christina L. Costantino

Contributed equally to this work with: Timothy K. Williams, Christina L. Costantino

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Nikolai A. Bildzukewicz

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Nathan G. Richards

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

David W. Rittenhouse

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Lisa Einstein

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Joseph A. Cozzitorto

Affiliation Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Judith C. Keen

Affiliation Robert Wood Johnson Medical School, University of Medicine and Dentistry of New Jersey, Camden, New Jersey, United States of America

Abhijit Dasgupta

Affiliations Department of Pathology, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Kimmel Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Myriam Gorospe

Affiliation Laboratory of Cellular and Molecular Biology, National Institute on Aging-Intramural Research Program, National Institutes of Health, Baltimore, Maryland, United States of America

Gregory E. Gonye

Affiliation Department of Pathology, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Charles J. Yeo

Affiliations Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Kimmel Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Agnieszka K. Witkiewicz

Affiliations Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Department of Pathology, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Kimmel Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Jonathan R. Brody

Jonathan.Brody@jefferson.edu

Affiliations Department of Surgery, Jefferson Pancreas, Biliary and Related Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Department of Pathology, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America, Kimmel Cancer Center, Thomas Jefferson University, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TW GG CLC AKW JCK. Performed the experiments: TW CLC NR DR NAB JCK JC AKW. Analyzed the data: TW CLC JC JRB AKW CJY NAB AD JCK LE MG NR DR GG. Contributed reagents/materials/analysis tools: CLC NAB TKW CJY JRB JAC LE AKW. Wrote the paper: JRB CJY TKW NAB CLC AKW JCK.