About the Authors

Nazanin S. Ruppender

Affiliations Department of Chemical and Biomolecular Engineering, Vanderbilt University, Nashville, Tennessee, United States of America, Center for Bone Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Alyssa R. Merkel

Affiliations Center for Bone Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Division of Clinical Pharmacology, Department of Medicine, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

T. John Martin

Affiliations Center for Bone Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Saint Vincent's Institute, Melbourne, Victoria, Australia

Gregory R. Mundy

† Deceased.

Affiliations Center for Bone Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Division of Clinical Pharmacology, Department of Medicine, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Veterans' Affairs Tennessee Valley Healthcare System, Nashville, Tennessee, United States of America

Julie A. Sterling

Affiliations Center for Bone Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Veterans' Affairs Tennessee Valley Healthcare System, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Scott A. Guelcher

scott.guelcher@vanderbilt.edu

Affiliations Department of Chemical and Biomolecular Engineering, Vanderbilt University, Nashville, Tennessee, United States of America, Center for Bone Biology, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NSR TJM GRM JAS SAG. Performed the experiments: NSR ARM. Analyzed the data: NSR JAS. Contributed reagents/materials/analysis tools: NSR SAG. Wrote the paper: NSR TJM JAS SAG.