About the Authors

Timothy Billings

Contributed equally to this work with: Timothy Billings, Evelyn E. Sargent

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Evelyn E. Sargent

Contributed equally to this work with: Timothy Billings, Evelyn E. Sargent

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Jin P. Szatkiewicz

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Nicole Leahy

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Il-Youp Kwak

Affiliation Department of Statistics, University of Wisconsin, Madison, Wisconsin, United States of America

Nazira Bektassova

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Michael Walker

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Terry Hassold

Affiliation Center for Reproductive Biology, Washington State University, Pullman, Wisconsin, United States of America

Joel H. Graber

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Karl W. Broman

Affiliation Department of Biostatistics & Medical Informatics, University of Wisconsin, Madison, Wisconsin, United States of America

Petko M. Petkov

Petko.Petkov@jax.org

Affiliation Center for Genome Dynamics, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PMP. Performed the experiments: TB EES TH. Analyzed the data: JPS NL IYK NB MW TH JHG KWB PMP. Contributed reagents/materials/analysis tools: NL NB MW JHG KWB. Wrote the paper: PMP.