About the Authors

Adam E. Handel

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Clinical Neurology, University of Oxford, Oxford, United Kingdom

Lynne Jarvis

Affiliation National Health Service Scotland, Edinburgh, United Kingdom

Ryan McLaughlin

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Clinical Neurology, University of Oxford, Oxford, United Kingdom

Anastasia Fries

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Clinical Neurology, University of Oxford, Oxford, United Kingdom

George C. Ebers

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Clinical Neurology, University of Oxford, Oxford, United Kingdom

Sreeram V. Ramagopalan

sreeramr@well.ox.ac.uk

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Clinical Neurology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AEH GCE SVR. Analyzed the data: AEH LJ RM AF SVR. Wrote the paper: AEH LJ RM AF GCE SVR.