About the Authors

Taiji Nakashima

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Masatoshi Jinnin

mjin@kumamoto-u.ac.jp

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Tomomi Etoh

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Satoshi Fukushima

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Shinichi Masuguchi

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Keishi Maruo

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Yuji Inoue

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Tsuyoshi Ishihara

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Hironobu Ihn

Affiliation Department of Dermatology and Plastic Surgery, Faculty of Life Sciences, Kumamoto University, Kumamoto, Japan

Competing Interests

This study was supported in part by grants from a commercial source (Rohto, Shiseido and Mitsubishi Pharma). The funders do not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TN. Performed the experiments: TN MJ TE SF SM KM YI TI HI. Analyzed the data: TN MJ TE SF SM KM YI TI HI. Wrote the paper: TN.