About the Authors

Jun Wei

Affiliation Division of Rheumatology, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Asish K. Ghosh

Current address: Feinberg Cardiovascular Research Institute, Northwestern University, Chicago, Illinois, United States of America.

Affiliation Division of Rheumatology, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Jennifer L. Sargent

Affiliation Department of Genetics, Dartmouth Medical College, Hanover, New Hampshire, United States of America

Kazuhiro Komura

Affiliation Division of Rheumatology, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Minghua Wu

Affiliation Division of Rheumatology, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Qi-Quan Huang

Affiliation Division of Rheumatology, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Manu Jain

Affiliation Division of Respiratory and Critical Care Medicine, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Michael L. Whitfield

Affiliation Department of Genetics, Dartmouth Medical College, Hanover, New Hampshire, United States of America

Carol Feghali-Bostwick

Affiliation Division of Pulmonary, Allergy, and Critical Care Medicine, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

John Varga

j-varga@northwestern.edu

Affiliation Division of Rheumatology, Northwestern University Feinberg School of Medicine, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JW AKG JV. Performed the experiments: JW AKG JLS KK MW MJ. Analyzed the data: JW AKG JLS MW MJ. Contributed reagents/materials/analysis tools: QQH MLW CFB JV. Wrote the paper: JW AKG JV.