About the Authors

Hannah L. Bader

Contributed equally to this work with: Hannah L. Bader, Alison L. Ruhe

Affiliation Department of Biomedical Engineering, Lerner Research Institute, Cleveland Clinic, Cleveland, Ohio, United States of America

Alison L. Ruhe

Contributed equally to this work with: Hannah L. Bader, Alison L. Ruhe

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Lauren W. Wang

Affiliation Department of Biomedical Engineering, Lerner Research Institute, Cleveland Clinic, Cleveland, Ohio, United States of America

Aaron K. Wong

Current address: Department of Computer Science, Princeton University, Princeton, New Jersey, United States of America

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Kari F. Walsh

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Rebecca A. Packer

Affiliation Department of Veterinary Clinical Sciences, Purdue University, West Lafayette, Indiana, United States of America

Jonathan Mitelman

Affiliation Kingston Road Animal Hospital, Toronto, Ontario, Canada

Kathryn R. Robertson

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Dennis P. O'Brien

Affiliation Department of Veterinary Medicine and Surgery, University of Missouri, Columbia, Missouri, United States of America

Karl W. Broman

Affiliation Department of Biostatistics and Medical Informatics, University of Wisconsin, Madison, Wisconsin, United States of America

G. Diane Shelton

Affiliation Department of Pathology, School of Medicine, University of California San Diego, La Jolla, California, United States of America

Suneel S. Apte

Affiliation Department of Biomedical Engineering, Lerner Research Institute, Cleveland Clinic, Cleveland, Ohio, United States of America

Mark W. Neff

mark.neff@vai.org

Current address: Translational Genomics Research Institute, Phoenix, Arizona, United States of America; The Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HLB ALR LWW SSA MWN. Performed the experiments: HLB ALR LWW AKW KFW RAP KRR DPO GDS. Analyzed the data: HLB ALR LWW AKW KFW RAP DPO KWB SSA MWN. Contributed reagents/materials/analysis tools: JM SSA. Wrote the paper: HLB LWW SSA MWN.